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Leaf beetle
The family Chrysomelidae, commonly known as leaf beetles, includes over 35,000 species in more than 2,500 genera, making it one of the largest and most commonly encountered of all beetle families. Numerous subfamilies are recognized, but only some of them are listed below. Leaf beetles are partially recognizable by their tarsal formula, which appears to be 4-4-4, but is actually 5-5-5.〔(【引用サイトリンク】 publisher = University of Florida )〕 Some lineages are only distinguished with difficulty from longhorn beetles (family Cerambycidae), namely by the antennae not arising from frontal tubercles. Adult and larval leaf beetles feed on all sorts of plant tissue. Many are serious pests of cultivated plants, for example the Colorado potato beetle (''Leptinotarsa decemlineata''), the asparagus beetle (''Crioceris asparagi''), the cereal leaf beetle (''Oulema melanopus''), and various flea beetles, and a few act as vectors of plant diseases. Others are beneficial due to their use in biocontrol of invasive weeds. Most Chrysomelidae are conspicuously colored, typically in glossy yellow to red or metallic blue-green hues, and some (especially Cassidinae) have spectacularly bizarre shapes. Thus, they are highly popular among insect collectors. == Description == The imagoes of leaf beetles are small to medium-sized, i.e. the size of their bodies varies from 1 to 18 millimeters. The bodies of most of their specimens are arched and egg-shaped, and they often possess a metallic luster or multiple colors. The head's shape (disregarding antennae and mouthparts) is roundish. In most specimens the antennae are notably shorter than head, thorax and abdomen, i.e. not more than half their combined length. The second antennae segment is of normal size (which differentiates leaf beatles from the closely related longhorn beetles). The antennae's segments are of a more or less equal shape, at most they gradually widen towards the tip. The first segment of the antennae, however, in most cases is larger than the following ones. Like the head, the eyes in most cases are also of a round shape. The pronotum of leaf beatles is arched in different shapes; only rarely it protrudes in bumpy shapes. The first three sternites are not fused; there are sutures. All leaf beetles possess wings. Only in some cases they are shortened, and they never leave more than the last tergum uncovered.
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